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PO Box 1346
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The Institute for Bird Populations
© 2002

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Peter Pyle (Staff Biologist) Tel: (415) 663 2053

Peter has been working for IBP since 1996. He attended Swarthmore College during the 1970's while also working seasonally on the Hawaii and other Pacific Forest Bird Surveys. He first worked for Dave DeSante in 1980 when he did internships for PRBO's Palomarin and Farallon Island Landbird Program, of which Dave was in charge. In 1985 Peter became a biologist on the Farallon Islands, a post he held until 2003. In 1996-2003 Peter split time between PRBO and IBP, and since 2003 he has been a full-time Biologist at IBP, doing scientific research, writing reports, and conducting banding workshops. Peter is also a Research Associate at the California Academy of Sciences in San Francisco and the Bishop Museum in Honolulu. Peter has authored or co-authored over 100 peer-reviewed papers, four books, and an on-line monograph of the birds of Hawaii. Among bird banders he is best known for his Identification Guide to North American Birds, Parts 1 and 2, which includes detailed criteria for ageing and sexing all North American birds in the hand and the field. In 2011 he had the good fortune of describing a new bird species, Bryan's Shearwater, and naming it after his grandfather.

Selected Publications

a. Senior Author

Pyle, P. 1975. Thirty-ninth breeding bird census: Mixed upland habitat, Montgomery County, Maryland. American Birds 29:11-15.

Pyle, P. 1976. Long-billed Curlew seen in Somerset County, Maryland. Maryland Birdlife 32:101.

Pyle, P., D.F. DeSante, R.J. Boekelheide, and R.P. Henderson. 1983. A Dusky Warbler on Southeast Farallon Island, California. Auk 100: 995-996.

Pyle, P. 1984. Observations of migrant and vagrant birds on Kure and Midway Atolls, 1982-1983. 'Elepaio 44:108-111.

Pyle, P. 1985. Identification Guide to European Passerines [Review]. North American Bird Bander 10:22-23.

Pyle, P., and J. Engbring. 1985. Checklist of the birds of Micronesia. 'Elepaio 46:57-68.

Pyle, P., and B. Eilerts. 1986. Pelagic seabird observations from Northwest Hawaiian Island waters. 'Elepaio 46:181-183.

Pyle, P., and W.C. Gagne. 1986. The globe skimmer, Pantala flavescens, on Kure Atoll. Proceedings of the Hawaii Entomological Society 26:4-5.

Pyle, P., and J. Engbring. 1987. New bird records and migrant observations from Micronesia, 1977-1984. 'Elepaio 47:11-15.

Pyle, P., S.N.G. Howell, R.P. Yunick, and D.F. DeSante. 1987. Identification Guide to North American Passerines. Slate Creek Press, Bolinas, CA. 280 pp.

Pyle, P. 1988. Comments and an errata for Identification Guide to North American Passerines. North American Bird Bander 13:112-113.

Pyle, P., and J. Engbring. 1988. First described nest and nest-site activity of the Truk Greater White-eye (Rukia ruki). Micronesica 21:281-283.

Pyle, P., P.V. Donaldson, R. David, and R.L. Pyle. 1988. The status of small Calidris sandpipers in the Hawaiian Islands: Documentation of three first state records. 'Elepaio 48:71-77.

Pyle, P., and P. Henderson. 1990. On separating female and immature Oporornis warblers in fall. Birding 22: 222-229.

Pyle, P., L. Spear, and J. Engbring. 1990. A previously unreported population of Herald Petrels on Ta'u Island, American Samoa. Colonial Waterbirds 13:136-138.

Pyle, P., and R.P. Henderson. 1991. The birds of Southeast Farallon Island: Occurrence and seasonal distribution of migratory species. Western Birds 22:41-84.

Pyle, P., L.B. Spear, W.J. Sydeman, and D.G. Ainley. 1991. The effects of experience and age on the breeding performance of Western Gulls. Auk 108: 25-33.

Pyle, P., and G. McCaskie. 1992. Thirteenth report of the California Bird Records Committee. Western Birds 23:97-132.

Pyle, P., and D.A. Sibley. 1992. Juvenal-plumaged Le Conte's Sparrows on migration. Birding 24:70-76

Pyle, P. 1993. A Markham's Storm-Petrel in the Northeastern Pacific. Western Birds 24:108-110.

Pyle, P., and S.N.G. Howell. 1993. An Arctic Warbler in Baja California, Mexico. Western Birds 24:53-56.

Pyle, P., and S.N.G. Howell. 1993. An aggregation of Peregrine Falcons on the Mexican winter grounds. Euphonia 2:92-94.

Pyle, P., L.B. Spear, and D.G. Ainley. 1993. Observations of Dark-rumped Petrels off Oregon and California. Western Birds 24:110-112.

Pyle, P., N. Nur, R.P. Henderson, and D.F. DeSante. 1993. The effects of weather and lunar cycle on nocturnal migration of landbirds at Southeast Farallon Island, California. Condor 95:343-361.

Pyle, P., and D.F. DeSante. 1994. Trends in waterbirds and raptors at Southeast Farallon Island, California, 1974-1993. Bird Populations 2:33-43.

Pyle, P., K. Hanni, and D. Smith. 1994. Bird notes from Isla Guadalupe, including three new island records. Euphonia 3:1-4.

Pyle, P., N. Nur, and D.F. DeSante. 1994. The landbird census on Southeast Farallon Island: An indicator of population change? Pp. 27-34 in S. Droege (ed.), Proceedings of the Migration Monitoring Workshop, U.S. Fish and Wildlife Service, Washington D.C.

Pyle, P., N. Nur, and D.F. DeSante. 1994. Trends in nocturnal migrant landbirds at Southeast Farallon Island, California. Studies in Avian Biology 15:58-74.

Pyle, P. 1995. Age of Norfolk Red-breasted Nuthatch. British Birds 88:611.

Pyle, P. 1995. Incomplete flight-feather molts and age in certain North American non-passerines. North American Bird Bander 20:15-26.

Pyle, P., and S.N.G. Howell. 1995. Flight-feather molt patterns and age in North American woodpeckers. Journal of Field Ornithology 66:564-581.

Pyle, P., and L. Gilbert. 1996. Occurrence patterns and trends of cetaceans recorded from Southeast Farallon Island, California, 1973-1994. Northwestern Naturalist 77:1-8.

Pyle, P., and S.N.G. Howell. 1996. Intraspecific variation and the identification of Spizella sparrows. Birding 28:374-387.

Pyle, P., S.D. Anderson, and D.G. Ainley. 1996. Trends in white shark predation at the South Farallon Islands, 1968-1993. Pp. 375-379 in A.P. Klimley and D.G. Ainley, eds., Great white sharks: The biology of Carcharodon carcharias. Academic Press, San Diego, CA.

Pyle, P., S.D. Anderson, A.P. Klimley, and R.P. Henderson. 1996. Environmental factors affecting the occurrence and behavior of white sharks at the Farallon Islands, California. Pp. 281-291 in A.P. Klimley and D.G. Ainley, eds., Great white sharks: The biology of Carcharodon carcharias. Academic Press, San Diego, CA.

Pyle, P. 1997. A further examination of wing and tail formulae in Contopus and Empidonax flycatchers. Pp. 147-154 in R.W. Dickerman, comp., The era of Allan Phillips: A Festschrift. R.W. Dickerman, Albequerque, NM.

Pyle, P. 1997. First year Slaty-backed Gull. Birders Journal 6:251.

Pyle, P. 1997. Flight-feather molt patterns and age in North American owls. Monographs in Avian Biology 2:1-32.

Pyle, P. 1997. Identification Guide to North American Birds. Part 1. Slate Creek Press, Bolinas, CA. 742 pp.

Pyle, P. 1997. Molt limits in North American passerines. North American Bird Bander 22:49-90.

Pyle, P., N. Nur, W.J. Sydeman, and S.D. Emslie. 1997. Cost of reproduction and the evolution of deferred breeding in the Western Gull. Behavioral Ecology 8:140-147.

Pyle, P., S.N.G. Howell, and G.M. Yanega. 1997. Molt, retained flight feathers and age in North American hummingbirds. Pp. 155-166 in R.W. Dickerman, comp., The era of Allan Phillips: A Festschrift. R.W. Dickerman, Albequerque, NM.

Pyle, P. 1998. A field guide to the warblers of North America [Review]. Western Birds 29:233-234.

Pyle, P. 1998. Eccentric first-year molts in certain Tyrannid flycatchers. Western Birds 29:29-35.

Pyle, P., and J. Nestler. 1998. First record of a Cuculus cuckoo on Midway Atoll and the Hawaiian Islands. Western Birds 29:124-127.

Pyle, P., and P. Unitt. 1998. Molt and plumage variation by age and sex in the California and Black-tailed gnatcatchers. Western Birds 29:280-289.

Pyle, P. 1999. Molts by age in the Bristle-thighed Curlew and other shorebirds. Western Birds 30:181-183.

Pyle, P., E. McLaren, and W.J. Sydeman. 1999. Organochlorine concentrations, eggshell thickness, and hatchability in seabirds off Central California. Colonial Waterbirds 22:376-381.

Pyle, P., M.J. Schramm, C. Keiper, and S.D. Anderson. 1999. Predation on a white shark (Carcharodon carcharias) by a killer whale (Orcinus orca) and a possible case of competitive displacement. Marine Mammal Science 15:563-568.

Pyle, P., and S.N.G. Howell. 2000. Revised ageing and sexing criteria for the Blue-throated Hummingbird. North American Bird Bander 25:134-137.

Pyle, P. 2001. Age at first breeding and natal dispersal in a declining population of Cassin's Auklet. Auk 118:996-1007.

Pyle, P. 2001. Isles of Refuge [Review]. Western Birds 32:234

Pyle, P., D.J. Long, J. Shonewald, R.E. Jones, and J. Roletto. 2001. Historical and recent colonization of the South Farallon Islands, California, by Northern Fur Seals (Callorhinus ursinus). Marine Mammal Science 17:397-402.

Pyle, P., N. Hoffman, B. Casler, and T. McKee. 2001. Little and Least terns breeding on Midway Atoll: Identification, range extensions, and assortative breeding behavior. North American Birds 55:3-6.

Pyle, P., W.J. Sydeman, and M. Hester. 2001. Effects of age, breeding experience, mate fidelity, and site fidelity on breeding performance in a declining population of Cassin's Auklets. Journal of Animal Ecology 70:1088-1097.

Pyle, P. 2002. Biological and Ecological Niches of the Gulf of the Farallones: Seabirds. Pp. 44-48 in K.A. Herman et al., eds., Beyond the Golden Gate-- oceanography, geology, biology, and environmental issues in the Gulf of the Farallones. USGS Circular

Pyle, P. 2002. Bird Station Manual [Review]. Auk 119:290-291.

Pyle, P., and D.F. DeSante. 2003. Four-letter and six-letter alpha codes for birds recorded in the American Ornithologists' Union Check-list area. North American Bird Bander 28:64-79.

Pyle, P. and S.N.G. Howell. 2004. Ornamental plume development and the "prealternate molts" of herons and egrets. Wilson Bulletin 116:287-297.

Pyle, P., A. Engilis Jr., and T.G. Moore. 2004. A Specimen of the nominate subspecies of the Red-Shouldered hawk from California. Western Birds 35:100-104

Pyle, P. 2005. First-cycle molts in North American Falconiformes. Raptor Research 39:378-385.

Pyle, P. 2005. Molts and plumages of ducks. Colonial Waterbirds 28:207-218.

Pyle, P. 2005. Remigial molt patterns in North American Falconiformes as related to age, sex, breeding status, and life-history strategies. Condor 107:823-834.

Pyle, P., and D.F. DeSante. 2005. Updates to four-letter and six-letter alpha codes based on revisions by the American Ornithologists' Union. North American Bird Bander 30:70-72.

Pyle, P. 2006. Handbook of avian hybrids of the world [Review]. Western Birds 37:237-239

Pyle, P. 2006. Offshore Pacific highlights in summer-fall 2005: Just another year. North American Birds 60:4-13.

Pyle, P. 2006. Staffelmauser and other adaptive wing-molt strategies in larger birds. Western Birds 37:179-185.

Pyle, P., and D.F. DeSante. 2006. Updates to four-letter and six-letter alpha codes based on revisions by the American Ornithologists' Union in 2005-2006. North American Bird Bander 31:193-195.

Pyle, P., G. Friedrichsen, T. Staudt, C. Oedekoven, and L.T. Ballance. 2006. First Record of Ringed Storm-Petrel (Oceanodroma hornbyi) for North America. North American Birds 60:162-163

Pyle, P. 2007. Revision of molt and plumage terminology in Ptarmigan (Phasianidae: Lagopus spp.) based on evolutionary considerations. Auk 124:508-514.

Pyle, P. 2008. Barefoot on Lava: The Journals and Correspondence of Naturalist R. C. L. Perkins in Hawai'i, 1892-1901 [Review]. Western Birds 39:230-232.

Pyle, P. 2008. Identification Guide to North American Birds. Part 2. Slate Creek Press, Point Reyes Station, CA.

P. Pyle. 2008. Birding by feather: A molt and plumage primer. Birding web extra. http://www.google.com/url?q=http://www.aba.org/birding/v40n6p18w1.pdf&sa=

Pyle, P., S.L. Jones, and J.M. Ruth 2008. Molt and aging criteria of four North American grassland passerines. Biological Technical Publication, FWS/BTP-R6011-2008. U.S. Fish and Wildlife Service, Washington, D.C.

Pyle, P. 2009. Age determination and molt strategies in North American alcids. Marine Ornithology 37:219-225.

Pyle, P., and D.F. DeSante. 2009. Updates to four-letter and six-letter alpha codes based on revisions by the American Ornithologists' Union in 2007-2008. North American Bird Bander 34:65-67.

Pyle, P., and D.F. DeSante. 2009. Updates to four-letter and six-letter alpha codes based on revisions by the American Ornithologists' Union in 2009. North American Bird Bander 34:109-110.

Pyle, P., W.A. Leitner, L. Lozano-Angulo, F. Avilez-Teran, H. Swanson, E. Gómez-Limón, and M.K. Chambers. 2009. Temporal, spatial, and annual variation in the occurrence of molt-migrant passerines in the Mexican monsoon region. Condor 111:583-590.

Pyle, P., and R. Kayhart. 2010. Replacement of primaries during the prealternate molt of a Yellow Warbler. North American Bird Bander 35:178-181.

Pyle, P., and B.L. Sullivan. 2010. Documenting repeated occurrence of individual birds with digital images. Western Birds: 41:261-265

Pyle, P. 2011. Nomenclature of the Laysan Honeycreeper, Himatione [sanguinea] fraithii. Bulletin of the British Ornithological Club 131:116-117.

Pyle, P. 2011. Paseriformes del Occidente de México: morfometría, datación y sexado. [Review]. Revista Catalana d'Ornitologia, L'Abellerol, and Western Birds 42:120-122.

Pyle, P., and M. B. Main. 2011. Surveying and monitoring birds. Pp. 49-90 in M.B. Main, ed., Wildlife monitoring: Florida Mastern Naturalist Program special topics. Southwest Florida Research and Education Center, Immokalee, FL. www.MasterNaturalist.org.

Pyle, P., J. Tietz, and G. McCaskie. 2011. Thirty-fifth report of the California Bird Records Committee: 2009 records. Western Birds 42:134-163.

Pyle, P. A.J. Welch, and R.C. Fleischer. 2011. A new species of shearwater (Puffinus) recorded from Midway Atoll, Northwestern Hawaiian Islands. Condor 113:518-527.

Pyle, P., D.L. Webster, and R.W. Baird. 2011. Notes on petrels of the Dark-rumped Petrel complex (Pterodroma phaeopygia/sandwichensis) in Hawaiian waters. North American Birds 65:364-367.

b. Coauthor

Ashman, P., and P. Pyle. 1978. First records of Lavender Fire Finch on Hawaii. 'Elepaio 38:99-100.

Ashman, P.R., P. Pyle, and J. Jeffrey. 1984. A second nest of the Small Kauai Thrush. 'Elepaio 45:33-34.

DeSante, D.F., and P.Pyle. 1986. Distributional Checklist to the Birds of North America. Artemesia Press, Lee Vining CA. 442 pp.

Howell, S.N.G., and P. Pyle. 1988. Additional notes on birds in Baja California. Aves Mexicanas 1:5-6.

Bailey, S.F., P. Pyle, and L. Spear. 1989. Dark Pterodroma petrels in the North Pacific: Identification, status, and Pacific North American occurrence. American Birds 43:400-415.

Howell, S.N.G., and P. Pyle. 1989. Another vagrant location in Baja California. Aves Mexicanas 2:3-4.

Howell, S.N.G., and P. Pyle. 1990. Additional notes on birds in Baja California, May 1989. Aves Mexicanas 2:6-7.

Sydeman, W.J., J.F. Penniman, T.M. Penniman, P. Pyle, and D.G. Ainley. 1991. Breeding performance in the Western Gull: Effects of parental age, timing of breeding and year in relation to food availability. Journal of Animal Ecology 60:135-149.

Emslie, S.D., W.J. Sydeman, and P. Pyle. 1992. The importance of mate retention and experience on breeding success in Cassin's Auklet (Ptychoramphus aleuticus). Behavioral Ecology 3:189-195.

Klimley, A.P., S.D. Anderson, P. Pyle, and R.P. Henderson. 1992. Spatiotemporal patterns of white shark (Carcharodon carcharias) predation at the South Farallon Islands, California. Copeia 1992:680-690.

Bartle, J.A., D. Hu, J.-C. Stahl, P. Pyle, T.R. Simons, and D. Woodby. 1993. Status and ecology of gadfly petrels in the temperate North Pacific. Pp. 101-111 in K. Vermeer, K.T. Briggs, K.H. Morgan, and D. Siegel-Causey, eds., The status, ecology, and conservation of marine birds of the North Pacific. Special Publication,
Howell, S.N.G., and P. Pyle. 1993. New and noteworthy bird records from Baja California, Mexico, October 1991. Western Birds 24:57-62.

Howell, S.N.G., P. Pyle, L.B. Spear, and R.L. Pitman. 1993. North American migrant birds on Clipperton Atoll. Western Birds 24:73-80.

Ralph, C.J., G.R. Geupel, P.Pyle, T.E. Martin, and D.F. DeSante. 1993. Handbook for field methods for monitoring landbirds. U.S. Department of Agriculture General Technical Report PSW-GTR-144, Albany, CA.

Howell, S.N.G., L.B. Spear, and P. Pyle. 1994. Identification of Manx-type shearwaters in the Eastern Pacific. Western Birds 25:169-177.

Ainley, D.G., R.L. Veit, S.G. Allen, L.B. Spear, and P. Pyle. 1995. Variations in marine bird communities of the California Current, 1986-1994. California Cooperative Oceanic Fisheries Investigations Reports 36:22-37.

Erickson, R.A., R.A. Hamilton, S.N.G. Howell, P. Pyle, and M.A. Patten. 1995. First record of the Marbled Murrelet and third record of the Ancient Murrelet for Mexico. Western Birds 26:39-45.

Spear, L., W.J. Sydeman, and P. Pyle. 1995. Factors affecting recruitment age and recruitment probability in the Western Gull Larus occidentalis. Ibis 137:352-359.

Anderson, S.D., A.P. Klimley, P. Pyle, and R.P. Henderson. 1996. Tidal height and white shark predation at the South Farallon Islands, California. Pp. 275-279 in A.P. Klimley and D.G. Ainley, eds., Great white sharks: The biology of Carcharodon carcharias. Academic Press, San Diego, CA.

Anderson, S.D., R.P. Henderson, P. Pyle, and D.G. Ainley. 1996. White shark reactions to unbaited decoys. Pp. 223-228 in A.P. Klimley and D.G. Ainley, eds., Great white sharks: The biology of Carcharodon carcharias. Academic Press, San Diego, CA.

Klimley, A.P., P. Pyle, and S.D. Anderson. 1996. Tail slap and breach: agonistic displays among white sharks? Pp. 241-255 in A.P. Klimley and D.G. Ainley, eds., Great white sharks: The biology of Carcharodon carcharias. Academic Press, San Diego, CA.

Klimley, A.P., P. Pyle, and S.D. Anderson. 1996. The behavior of white sharks and their pinniped prey during predatory attacks. Pp. 175-191 in A.P. Klimley and D.G. Ainley, eds., Great white sharks: The biology of Carcharodon carcharias. Academic Press, San Diego, CA.

Long, D.J, K. Hanni, P. Pyle, J. Roletto, R.E. Jones, and R. Bandar. 1996. White shark predation on four pinniped species in central California waters: geographical and temporal patterns inferred from wounded carcasses. Pp. 263-274 in A.P. Klimley and D.G. Ainley, eds., Great white sharks: The biology of Carcharodon carcharias. Academic Press, San Diego, CA.

Ralph, C. J., G. R. Geupel, P. Pyle, T. E. Martin, D. F. DeSante, and B. Mila. 1996. Manual de Métodos de Campo Para el Monitoreo de Aves Terrestres. USDA Forest Service, General Technical Report PSWGTR-159. Pacific Southwest Research Station, Albany, CA.

Sydeman, W.J., P. Pyle, S.D. Emslie, and E.B. Mclaren. 1996. Causes and consequences of long-term partnerships in Cassin's Auklets. Pp. 211-222 in J.M. Black (ed.), Partnerships in Birds. Oxford University Press, U.K.

Veit, R.R., P. Pyle, and J.A. McGowan. 1996. Ocean warming and long-term change of pelagic bird abundance within the California Current System. Marine Ecology Progress Series 139:11-18.

Hanni, K.D., D.J. Long, R.E. Jones, P. Pyle, and L.E. Morgan. 1997. Sightings and strandings of Guadalupe fur seals in central and northern California. Journal of Mammalogy 78:684-690.

Howell, S.N.G., and P. Pyle. 1997. Twentieth report of the California Bird Records Committee: 1994 records. Western Birds 28:117-141.

Sydeman, W.J., K.A. Hobson, P. Pyle, and E.B. McLaren. 1997. Trophic relationships among seabirds in Central California: Combined stable isotope and conventional dietary approach. Condor 99:327-336.

Veit, R.R., J.A. McGowan, D.G. Ainley, T.R. Wahl, and P. Pyle. 1997. Apex marine predator declines ninety percent in association with changing oceanic climate. Global Change Biology 3:23-28.

Yanega, G.M., P. Pyle, and G.R. Geupel. 1997. The timing and reliability of bill corrugations for ageing hummingbirds. Western Birds 28:13-18.

Mills, K.L., P. Pyle, W.J. Sydeman, J. Buffa, and M.J. Rauzon. 2002. Direct and indirect effects of house mice on declining populations of a small seabird, the ashy storm-petrel (Oceanodroma homochroa) on Southeast Farallon Island, California. P. 412 in C. R. Veitch and N. M. Clout (eds), Turning the Tide: the Eradication of Invasive Species: Proceedings of the International Conference on Eradication of Island Invasives, Gland, Switzerland

Nur, N., W. J. Sydeman, M. Hester, and P. Pyle. 1998. Survival in Cassin's Auklet on Southeast Farallon Island: Temporal patterns, population variability, and the cost of double-brooding. Pacific Seabirds 25:38-39

Spear, L.B., P. Pyle, and N. Nur. 1998. Natal dispersal in the Western Gull: Proximal factors and fitness consequences. Journal of Animal Ecology 67:165-179.

DeSante, D.F., D.R. O'Grady, and P. Pyle. 1999. Measures of productivity and survival derived from standardized mist-netting are consistent with observed population changes. Bird Study 46:S178-188.

Gochfeld, M., D.J. Gochfeld, D. Minton, B.M. Murray Jr., P. Pyle, N. Seto, and J. Burger. 1999. Metals in feathers of Bonin Petrel, Christmas Shearwater, Wedge-tailed Shearwater, and Red-tailed Tropicbird in the Hawaiian Islands, Northern Pacific Ocean. Environmental Monitoring and Assessment 59:343-358.

Heindel, M., and P. Pyle. 1999. Separating Western and Yellow-bellied Flycatchers in the field. Birders Journal 8:78-87.

Spear, L.B., D.G. Ainley, and P. Pyle. 1999. Seabirds in southeastern Hawaiian waters. Western Birds 30:1-32.

Capitolo, P., W. Richardson, R. Burnett, and P. Pyle. 2000. First record of an Olive-backed Pipit in California. Western Birds 31:112-116.

Hanni, K.D., and P. Pyle. 2000. Entanglement of pinnipeds in synthetic materials at Southeast Farallon Island, California, 1976-1998. Marine Pollution Bulletin 40:1076-1081.

Abbott, S, S.N.G. Howell, and P. Pyle. 2001. First North American Record of Greater Sandplover. North American Birds 55:252-257.

Earnheart-Gold, S., and P. Pyle. 2001. Occurrence patterns of Peregrine Falcons on Southeast Farallon Island, California, by subspecies, age, and sex. Western Birds 32:119-126.

Boustany, A.M., S.F. Davis, P. Pyle, S.D. Anderson, B.J. Le Boeuf, and B.A. Block. 2002. Expanded ecological niche for white sharks. Nature 415:35-36.

Howell, S.N.G., and P. Pyle. 2002. Ageing and molt in non-breeding Black-bellied Plovers. Western Birds 33:268-270

McKee, T. and P. Pyle. 2002. Plumage variation and hybridization in Black-footed and Laysan albatrosses. North American Birds 56:131-138.

Nott, M.P., DeSante, D.F., Siegel, R.B., and P. Pyle. 2002. Influences of the El Niño/Southern Oscillation and the North Atlantic Oscillation on avian productivity in forests of the Pacific Northwest of North America. Global Ecology and Biogeography 11:333-342.

Richardson, T.W., P. Pyle, R. Burnett, and P. Capitolo. 2003. The Occurrence and seasonal distribution of migratory birds on Southeast Farallon Island, 1968-1999. Western Birds 34:58-96.

Howell, S.N.G. and P. Pyle 2005. Molt, age determination, and identification of puffins. Birding 37:412-418.

LeValley, R., and P. Pyle. 2007. Notes on plumage maturation in the Red-tailed Tropicbird. Western Birds 38:306-310.

Weng, K.C., A. M. Boustany, P. Pyle, S. D. Anderson, A, Brown, and B. A. Block. 2007. Migration and habitat of White Sharks (Carcharodon carcharias) in the eastern Pacific Ocean. Marine Biology 152:857-894.

Pagen, R., P. Pyle, and L.T. Ballance. 2008. A Fregetta Storm-Petrel off Western Mexico. Western Birds 39:225-227.

Pyle, R.L., and P. Pyle. 2009. The Birds of the Hawaiian Islands: Occurrence, history, distribution, and status. B.P. Bishop Museum, Honolulu, HI, U.S.A. Version 1 (31 December 2009) http://hbs.bishopmuseum.org/birds/rlp-monograph

Wolfe, J.D., P. Pyle, and C.J. Ralph. 2009. Breeding seasons, molt patterns, and gender and age criteria for selected northeastern Costa Rican resident landbirds. Wilson Journal of Ornithology 121:556-567.

Chandler, R.M., P. Pyle, M.E. Flannery, D.J. Long, and S.N.G. Howell. 2010. Flight feather molt of Turkey Vultures. Wilson Journal of Ornithology 122:354-360.

Wolfe, J.D., T.B. Ryder, and P. Pyle. 2010. Using molt cycles to categorize age in tropical birds: An integrative system. Journal of Field Ornithology 81:186-194.

Chambers, M., G. David, C. Ray, B. Leitner, and P. Pyle. 2011. Habitats and conservation of molt-migrant birds in southeastern Arizona. The Southwestern Naturalist 56:204–211.

Radley, P., A.L. Crary, J. Bradley, C. Carter, and P. Pyle. 2011. Molt patterns, Biometrics, and age and gender classification of landbirds on Saipan, Northern Mariana Islands. Wilson Journal of Ornithology 123:588-594.

Wolfe, J.D., and P. Pyle. 2011. First evidence for eccentric prealternate molt in the Indigo Bunting (Passerina cyanea): Possible implications for adaptive molt strategies. Western Birds: In press.

Junda, J., A.L. Crary, and P. Pyle. In press. Two modes of primary replacement during prebasic molt of Rufous Fantails Rhipidura rufifrons. Journal of Field Ornithology.

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